DIGITALNA ARHIVA ŠUMARSKOG LISTA
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ŠUMARSKI LIST 1-2/2009 str. 63 <-- 63 --> PDF |
I. Dakskobler: FITOCENOLOŠKAISTRAŽIVANJA ŠUMSKIH EKOSUSTAVANA POČETKU 21. STOLJEĆAŠumarski list br. 1–2, CXXXIII (2009), 53-62 Society for Vegetation Ecology in the 1970s and 1980s, was also a map of natural potential vegetation of Yugoslavia in the scale of 1:1.000.000 (B. Jovanović et al. 1986) and Prodromus phytocoenosum Jugoslaviae (Zupančič et al. 1986). The work of the time was incorporated also into the Map of Natural Vegetation of Europe in the scale of 1:2500000 (Bohn et al. 2000). Development of fast and more advanced personal computers in the 1980 s, which paved a way for relatively simple massive utilisation of mathematical methods (above all multivariate statistics) in comparisons of phytocoenological relevés and their grouping by environmental factors, brought about a significant turnaround in vegetation research conducted according to the Central-European and other methods. One of the first widely used software of this kind was TWINSPAN (Hill 1979). Later on other program packages, such as MULVA (Wildi & Orloci 1996), SYN-TAX (Podani 2001), JUICE (Tichy2002), CANOCO (Ter Braak & Šmilaure 2002), PC-ORD (McCune & Mefford 2006), etc. were applied as well. In this respect, a problematic issue in the Central-European method is the subjective selection of relevés and subjective evaluation of cover or abundance of species with ordinal values (e.g. r, +, 1, 2, 3, 4, 5). There have been discussions among experts on the correct procedures for numeric processing of ordinal input data. Some, e.g. Podani (2005), believe that only ordinal classification and non-metric ordination methods are suitable for such data. Others disagree. A similar problem exists with the statistical analysis of data acquired using non-random (subjective) sampling, such as are also our relevés. Experts published their pro et contra views on when and to what extent such analysis is appropriate in the journal Folia geobotanica (Herben & Chytrý 2007). Despite the above concerns it is still true that the Central-European method allows a relatively fast, simple and inexpensive way of acquiring useful data on vegetation and its connections with the environment. Databases of vegetation relevés (e.g. TURBOVEG – Hennekens & Schaminée 2001) already keep large amounts of historic, several decades and even half a century old relevés that were made with subjective plot selection. Disregarding these relevés on account of their statistically problematic (subjective and non-random) origin would mean discarding very valuable ecological data. Ecologists therefore use these data to their advantage, but with regard to their limitations. These data are used also in contemporary overviews of vegetation of large regions (e.g. in Willner & Grabherr 2007). Using and processing large quantities of relevés has changed the views of the basic unit of the syntaxonomic system – association –in many ways, and has affected the way we see the concept of character and differential species (comp. Willner 2006). When selecting diagnostic species authors apply different computing procedures. A large number of relevés enable a relatively objective calculation of fidelity of species to certain syntaxa and their diagnostic value (e.g. with phi-coefficient – Tichý & Chytrý 2006). As a rule, in formalized classification the number of syntaxonomic units of a vegetation formation (e.g. forest communities) within a certain region is reduced. The question remains, however, whether such reduction is founded on the actual site conditions and on the actual phytocoenoses. Before the turn of the century there was a shift from the knowledge (study) of plant communities to the knowledge (study) of habitats. It is an acknowledgement of the Braun-Blanquet method that the most widely used habitat type classification in Europe (Devillers & J. Devillers-Teschuren 1996) is in many ways based on this method itself, as well as on its findings and its review of plant communities, arranged in a hierarchical system. If we compare Braun-Blanquet’s Phytocoenology from 1964 and van der Maarel’s Vegetation ecology, which was published in 2005, we can observe a significant development and a broad array of different approaches to the |