DIGITALNA ARHIVA ŠUMARSKOG LISTA
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ŠUMARSKI LIST 11-12/2010 str. 33 <-- 33 --> PDF |
M. Cojzer, R. Brus: SPECIES COMPOSITION AND SUCCESSIONAL PATHWAYS ON ABANDONED ... Šumarski list br. 11–12, CXXXIV (2010), 581-591 sing plots were the size, the shape and the developmental stage of the abandoned land. All plots were square shaped and north-east oriented. Vegetation records were carried out on 52 plots in total. 37 plots (20x20 m) were set on abandoned agricultural land, of which 5 were in young growth developmental stages, 15 in a thicket phase, and 17 in a pole stand phase. 15 plots (20x10 m), 5 for each developmental phase, were analysed in forest areas. The survey was carried out from the beginning of August until the end of October 2006, and in September 2007. Each plot was set according to its developmental phase (one plot – one developmental phase), and its floristic composition was recorded by counting the individuals of tree and shrub species. The source of the plant nomenclature was the Mala flora Slovenije (Martinčič et al., 2007). To analyse the vegetation composition for every single developmental phase, an appropriate treatment was assigned to each study plot. Six equivalent treatments were: A – abandoned land in young growth phase, B – abandoned land in thicket phase, C – abandoned land in pole stand phase, D – forest in young growth phase, E – forest in thicket phase, F – forest in pole stand phase. All vegetation samples which were carried out on abandoned land, together created Z treatment, and all vegetation samples which were carried out on forest plots, formed G treatment. 2. 3 Statistical analyses – Statističke analize MapInfo v. 8.5 software was used for spatial data processing, and Excel for data analysis. Statgraphics Plus for Windows software was used for statistical data processing. Data were analysed by analysis of variance (One-way ANOVA), where parametric (t-test) and nonparametric tests (LSD test, Duncan variance homogeneity test) were used. Since the number of subjects per ha is not a normally distributed variable, a preliminary »square root« transformation was performed for the ANOVA. Species diversity was calculated by the Shannon’s diversity index (H´; H´= -.(pi ln pi)) (Shannon, 1948). CANOCO 4.5 for Windows (ter Braak and Šmilauer, 2002) was used for calculation. The number of species per plot (species composition), their stability, and species density (the number of species individuals per ha) were calculated for each plot separately. Species stability was calculated according to the share of individual species occurrence in the previous samples. We analysed the plot similarity vegetation composition by using the detrended correspondence analysis (DCA), considering the number of individuals per ha. DCA analysis was carried out with the help of CANOCO 4.5 for Windows (ter Braak and Šmilauer, 2002). 3 RESULTS – Rezultati 3. 1 The share of forest area and abandoned land – Udio šumskih površina i zemljišta u zarastanju In twenty years (1985-2005) the forest area of Haloze increased by 6.9 % (Tab. 1). The share of forest cover in the Haloze region is thereby 42.2 %, but twenty years ago it used to be 39.4 %. And, if (by the next forest management plan renewal) all presently recorded areas of abandoned land are classified as forest, in 2015 (comparably to the year 1985) the forest area will have increased by 13.0 %, and the total forest area of Haloze will total 44.5 %. Table 1 Changes in the forest area in Haloze in the period from 1985 to 2005 (ZGS, 2005) Tablica 1. Promjena šumskih površina u Halozama od 1985 do 2005 godine Year Godina Forest area Površina šuma (ha) Index Indeks (%) 1985 6916.70 100.0 2005 7396.70 106.9 2015 7803.49 112.8 3. 2 Species composition and density of individuals (on abandoned land and forest) – Vrste drveća i grmlja te gustoća jedinki (na napuštenom zemljištu te u šumi) Abandoned land was more species diverse than forest; 47 species were identified on abandoned land (33 tree and 14 shrub species), and 36 species in forest (25 tree and 11 shrub species) (Tab. 4). With regard to the treatment, the ordination shows (Fig. 2), that by the first axis, which explains most of the variability, we can distinguish between two groups of samples: abandoned land (Z – treatment) and forest (G - treatment). The total inertion of DCA is 6.281; the eigenvalues of the first four axes were 0.728, 0.470, 0.322, 0.201, which cumulatively explained 11.6 %, 19.1 %, 24.2 %, 27.4 % of variability in species data. Gradient length of the first axis is 4.663, which justifies the use of uni-modal ordination methods (ter Braak and Šmilauer, 2002). The second axis shows a relatively high variability in vegetation composition per plot on abandoned land and a little lower variability in forest. Table 2 shows the average number of species per plot. There was no difference in average number of all species per plot, and in the average number of tree spe |