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ŠUMARSKI LIST 3-4/2011 str. 50     <-- 50 -->        PDF

Z. Sedlar, V. Hršak, R. Šoštarić: NUMERICALAND PHYTOSOCIOLOGICALANALYSIS OF ... Šumarski list br. 3–4, CXXXV (2011), 1-3


own special endemic taxon fromPinus nigras.l. complex
and can be rich in endemics (Brullo &al. 2001).
The size of the Mediterranean basin and the geographical
isolation of its islands and peninsulas drove to massive
speciation process among the species forming in the
high mountain vegetation.The biodiversity of Mediterranean
orophilous communities is thus among the highest
in Europe.This is why many local species enter into
speces assemblage of syntaxa of eachPinus nigrasubspecies,
as do species from neighbouring vegetation
type. In the Junipero sibiricae -Pinetum dalmaticae
Domac (1956) 1965 association species enter most
frequently from Quercetalia pubescentis Br.Bl. (1931)
1932 order. Black pine forests often have in their structure
a great number of species which grow on screes and
rocks than zonal vegetation (Bergmeier 1990). This
is why communities of different Mediterranean areas,
formed by black pine, have low number of matching
species and more expressed local floristic features than
it is the case with zonal and other types of vegetation
(Bergmeier 1990).The syntaxonomic classification
is very difficult and an object of different discussions
and different syntaxonomic solutions because of these
reasons (Rivas-Martínez 1964; Domac 1965;
Horvat &al. 1974; Trinajstić 1977, 1986, 1990,
1998; Bergmeier 1990; 2002; Rivas-Martínez
&al. 1999;Brullo&al.2001).


In classification of black pine forests there are several
different concepts. The first concept is taking a
pure numeric proportion of species from different syntaxa
with which black pine forests adjoins as a classification
basis for a higher level of syntaxa.According to
this concept it is attempted to classify black pine
forests into an existing higher level syntaxon or to present
it as a subassociation or a facies of a neighbouring
association which has the most matching species in
species assemblage. The second concept introduces
habitat differentiation, ecological and structural characteristics
which make numeric part of different syntaxa
relative. The third concept is to give greater
meaning to the supraregional characteristics, of termophillous
pine forests at merging to higher level syntaxa
(Bergmeier 1990). Example for this concept is
Erico-Pinetea Horvat 1959 class and lower syntaxa
(Horvat 1959).


Pinus nigrasubsp.dalmatica(Vis) Franco is an endemic
taxon distributed on the coast, on some islands
of southern Dalmatia and on the southern slopes of the
DinaricAlps at altitudes from 350 to 1,000 m and is
one of the Mediterranean taxa of the Pinus nigra s.l.
complex (Bogunić &al. 2003;Isajev &al. 2004;
Pignatti 2005). It is a woody phanerophyte of the
northeastern eurimediterranean, found in drier areas
(Pignatti 2005). It is also an Illyrian floral element
(Pignatti 1982a).


The distribution area of Dalmatian pine is spread
over altitudes between 300 and 1,400m and is divided
into two parts: the Biokovo Mountain part and the
coastal part (Trinajstić1986). Dalmatian pine vegetation
is mostly found in forms of different stages of
degradation ofQuercus ilexL. vegetation, mostly garrigues
(Stefanović 1977).The largest complexes of
Dalmatian pine vegetation are on the islands of Brač,
Hvar, and Korčula and on the Pelješac peninsula (Figure
1), at altitudes between 300 and 750 metres. Here,
they are situated in the Mediterranean region in both
eumediterranean and submediterranean vegetation zo nes.
Dalmatian pine vegetation is situated on Biokovo
Mountain mostly in the northwestern part of the mountain
and is divided into two altitudinal parts: an upper
part at an altitude of 1,100-1,400 m, and a lower one at
an altitude of 500-900 m (Domac 1961/62).


Until now, researchers have encountered problems
connected to the typology of Dalmatian pine vegetation.
It was often described as a unique type of vegetation
(Beck-Mannagetta 1901; Horvatić 1928;
Horvat 1950, 1954). Horvat 1950 and 1954 even
marked it as a separate association with its temporary
name “Pinetum dalmaticae”, but made no further attempt
at syntaxonomical analysis.


Later,Horvatić 1958 defined Dalmatian pine
vegetation only as the degradation stages of different
associations and gave them a subassociation and facies
level (Genisto-Ericetum verticillatae pinetosum dalmaticaeHorvatić
1958,Erico-Rosmarinetum pinetosum
dalmaticae Horvatić 1958, Stipo-Salvietum
officinalis pinetosum dalmaticae Horvatić 1958,
Orno-Quercetum ilicis pinosum dalmaticae Horvatić1958).


In contrast, Trinajstić 1977 named a new, higher
level,syntaxa, found on the island of Hvar and des cribed
as Erico manipuliflorae-Pinetum dalmaticae
Trinajstić 1977 and in 1986 another association from
the islands of Korčula and Brač,Querco ilicis-Pinetum
dalmaticaeTrinajstić 1986.


The research and classifications of Horvatić 1958
andTrinajstić 1977, 1986 and 1990 do not include the
area of Biokovo Mountain.The only attempt to describe
and classify Dalmatian pine vegetation on Biokovo
Mountain was made by Domac in 1965. He takes the
syntaxa from Horvatić 1958 into consideration and describes
a new association, found on the higher part of
Biokovo Mountain, asJunipero sibiricae -Pinetum dalmaticaeDomac1965,
classifying it into the deciduous
submediterranean Quercetalia pubescentis order, the
Querco-FageteaBr.-Bl. et Vlieger 1937 class.


This review shows that the syntaxonomy of the Dalmatian
pine vegetation is not simple, although it takes
up a relatively small area. It shows that there are no