DIGITALNA ARHIVA ŠUMARSKOG LISTA
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ŠUMARSKI LIST 3-4/2011 str. 56 <-- 56 --> PDF |
Z. Sedlar, V. Hršak, R. Šoštarić: NUMERICALAND PHYTOSOCIOLOGICALANALYSIS OF ... Šumarski list br. 3–4, CXXXV (2011), 1-3 and theCarpinion orientalisHorvat1958 alliance from Querco-Fagetea class, based on presence of more species from these syntaxa. However species from this order are present here with a low constancy. Further more,Junipero sibiricae -Pinetum dalmaticae is not a deciduous forest, it does not have deciduous tree species, and bushes are present with a low constancy. This association is rather dwarf shrub-like vegetation dominated by Dalmatian pine. Domac1965 specifiesPinus nigrasubsp.dalmatica and Sesleria robusta as characteristic species and Juniperus communis L. ssp. nana,Cerastium grandiflorum Waldst. & Kit. and Cynanchum adriaticum Beck (Fritsch) as differential species to differentiate this vegetation from others containing Pinus nigra subsp.dalmatica.All these species, except Dalmatian pine, have a high indicator value (IV) for the specific cluster in our analysis. In contrast,Pinus nigrasubsp. dalmaticadid not have any statistically significant indicator value because of its presence on both the coastal part and on Biokovo Mountain.Therefore, Dalmatian pine cannot be taken into account as a characteristic species for the Junipero sibiricae -Pinetum dalmaticaeassociation. Trinajstić 1986 had a different approach. He classified Junipero sibiricae-Pinetum dalmaticae into Erico-Pineteaclass. Horvat 1958, while defining this class, made species composition more relative, by using mutual supraregional characteristics.Therefore he indicated as class’characteristic species ones also characteristic for other termophillous syntaxa. This way there are as much arguments for classifying Junipero sibiricae-Pinetum dalmaticae into Erico- Pinetea and Erico-Pinetalia as there are for Quercetalia pubescentis. Erico-Pinetea according to its concept comprises termophillous coniferous forests, but in a more continental areas without expressed summer drought, which is present near the coast where Biokovo mountain is situated. Similar concept represents Bergmeier 1990 who classified Seslerio robustae- Pinetum pallasianaealso intoErico-Pinetea. The ISA showed species of different distribution areas as indicator species.Juniperus communisL. ssp. nana reveals a eurasiatic distribution, and, in this study, is determined as one of the two most important indicator species. Several indicator species show rather montane and alpine conditions present in Junipero sibiricae-Pinetum dalmaticae habitats. Together with these species, there are three species of endemic chorotype (Pignatti 2005):Sesleria robusta, defined by Domac 1965 as characteristic, and Cynanchum adriaticum and Cerastium grandiflorum as differential species for this association. All three species are distributed on the coastal part of Croatia and DinaricAlps.Sesleria robustais, with the same indicator value (IV) and probability (p) value asJuniperus communisL. ssp.nana, the second most indicative species. Out of these two species, Sesleria robusta shows Mediterranean, andJuniperus communisL. ssp. nanaalpine characteristics.S. robustais a species of an endemic chorotype (Pignatti 2005). It is distributed in the central Mediterranean (Pignatti 1982b), and in central and southern Dalmatia (FCD 2007), which gives a Mediterranean ecological character to this association. On the other hand,Juniperus communis L. ssp. nana is an alpine species (Vidaković 1982).This suggests that the habitats ofJunipero sibiricae -Pinetum dalmaticae have both Mediterranean and alpine ecological characteristics on Biokovo Mountain. Based on these species and their characteristics, we decided to carry out a new approach for the syntaxonomical classification of Junipero sibiricae- Pinetum dalmaticae. In spite the fact there are not many floristic arguments, but accepting the concept of mutual ecological, bio geographical and structural characteristics, we believe that Junipero sibiricae-Pinetum dalmaticae should not be put inQuercetalia pubescentis, nor into Erico-Pinetea, but into Pino-Juniperetea class. This class was first described byRivas-Martínez 1964 in west Mediterranean and it comprises a group of orophilous woody communities dominated by gymnosperms occurring in Spain.The structure of this vegetation is generally consisted by tree layer with more or less open canopies, thick shrub layer, and rather poor species assemblage. It also has relic character (Pignatti 1985, 1998;Brullo &al. 2001). Nowadays more associationsare describedfromPino-Juni pe reteaclass,scattered throughout the Me diterranean area.Vegetation belonging to this class is on arid soil and is a final stage of climatogenous vegetation for southern Eurosiberian and Mediterranean areas (Rivas- Martínez &al.1991).Pino-Junipereteaclass consists of orophilous communities dominated by conifers and its range extends to the whole Mediterranean basin.Brullo &al. 2001have expanded the distribution area of thePino-Junipereteainto eastern Mediterranean describing the eastern Mediterranean alliance Berberido creticae-Juniperion foetidissimae Brullo & al. 2001 which is distributed in Greece, Cyprus and western and southern Anatolia. This alliance belongs to Juniperetalia haemisphaericae Ri vas- Martínez &Molina 1999 order distributed in central and eastern Mediterranean. Within the Pino-Juniperetea class the pioneer orophilous vegetation counts, in general, quite a low number of specialized species. In spite of this, the lack of character species is compensated by their high biogeographic significance and allegiance to a precise ecological context.Pino-Junipereteacommunities ac |