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ŠUMARSKI LIST 3-4/2011 str. 56     <-- 56 -->        PDF

Z. Sedlar, V. Hršak, R. Šoštarić: NUMERICALAND PHYTOSOCIOLOGICALANALYSIS OF ... Šumarski list br. 3–4, CXXXV (2011), 1-3


and theCarpinion orientalisHorvat1958 alliance from
Querco-Fagetea class, based on presence of more
species from these syntaxa. However species from this
order are present here with a low constancy. Further
more,Junipero sibiricae -Pinetum dalmaticae is not a
deciduous forest, it does not have deciduous tree
species, and bushes are present with a low constancy.
This association is rather dwarf shrub-like vegetation
dominated by Dalmatian pine.


Domac1965 specifiesPinus nigrasubsp.dalmatica
and Sesleria robusta as characteristic species and
Juniperus communis L. ssp. nana,Cerastium grandiflorum
Waldst. & Kit. and Cynanchum adriaticum
Beck (Fritsch) as differential species to differentiate
this vegetation from others containing Pinus nigra
subsp.dalmatica.All these species, except Dalmatian
pine, have a high indicator value (IV) for the specific
cluster in our analysis. In contrast,Pinus nigrasubsp.
dalmaticadid not have any statistically significant indicator
value because of its presence on both the
coastal part and on Biokovo Mountain.Therefore, Dalmatian
pine cannot be taken into account as a characteristic
species for the Junipero sibiricae -Pinetum
dalmaticaeassociation.


Trinajstić 1986 had a different approach. He
classified Junipero sibiricae-Pinetum dalmaticae into
Erico-Pineteaclass. Horvat 1958, while defining this
class, made species composition more relative, by
using mutual supraregional characteristics.Therefore
he indicated as class’characteristic species ones also
characteristic for other termophillous syntaxa. This
way there are as much arguments for classifying
Junipero sibiricae-Pinetum dalmaticae into Erico-
Pinetea and Erico-Pinetalia as there are for Quercetalia
pubescentis. Erico-Pinetea according to its
concept comprises termophillous coniferous forests,
but in a more continental areas without expressed summer
drought, which is present near the coast where
Biokovo mountain is situated. Similar concept represents
Bergmeier 1990 who classified Seslerio robustae-
Pinetum pallasianaealso intoErico-Pinetea.


The ISA showed species of different distribution
areas as indicator species.Juniperus communisL. ssp.
nana reveals a eurasiatic distribution, and, in this
study, is determined as one of the two most important
indicator species. Several indicator species show
rather montane and alpine conditions present in
Junipero sibiricae-Pinetum dalmaticae habitats. Together
with these species, there are three species of endemic
chorotype (Pignatti 2005):Sesleria robusta,
defined by Domac 1965 as characteristic, and Cynanchum
adriaticum and Cerastium grandiflorum as
differential species for this association. All three
species are distributed on the coastal part of Croatia
and DinaricAlps.Sesleria robustais, with the same indicator
value (IV) and probability (p) value asJuniperus
communisL. ssp.nana, the second most indicative
species. Out of these two species, Sesleria robusta
shows Mediterranean, andJuniperus communisL. ssp.
nanaalpine characteristics.S. robustais a species of
an endemic chorotype (Pignatti 2005). It is distributed
in the central Mediterranean (Pignatti 1982b),
and in central and southern Dalmatia (FCD 2007),
which gives a Mediterranean ecological character to
this association. On the other hand,Juniperus communis
L. ssp. nana is an alpine species (Vidaković
1982).This suggests that the habitats ofJunipero sibiricae
-Pinetum dalmaticae have both Mediterranean
and alpine ecological characteristics on Biokovo
Mountain. Based on these species and their characteristics,
we decided to carry out a new approach for the
syntaxonomical classification of Junipero sibiricae-
Pinetum dalmaticae.


In spite the fact there are not many floristic arguments,
but accepting the concept of mutual ecological,
bio geographical and structural characteristics, we believe
that Junipero sibiricae-Pinetum dalmaticae
should not be put inQuercetalia pubescentis, nor into
Erico-Pinetea, but into Pino-Juniperetea class. This
class was first described byRivas-Martínez 1964
in west Mediterranean and it comprises a group of
orophilous woody communities dominated by gymnosperms
occurring in Spain.The structure of this vegetation
is generally consisted by tree layer with more
or less open canopies, thick shrub layer, and rather
poor species assemblage. It also has relic character
(Pignatti 1985, 1998;Brullo &al. 2001). Nowadays
more associationsare describedfromPino-Juni pe
reteaclass,scattered throughout the Me diterranean
area.Vegetation belonging to this class is on arid soil
and is a final stage of climatogenous vegetation for
southern Eurosiberian and Mediterranean areas (Rivas-
Martínez &al.1991).Pino-Junipereteaclass
consists of orophilous communities dominated by
conifers and its range extends to the whole Mediterranean
basin.Brullo &al. 2001have expanded the
distribution area of thePino-Junipereteainto eastern
Mediterranean describing the eastern Mediterranean
alliance Berberido creticae-Juniperion foetidissimae
Brullo & al. 2001 which is distributed in Greece,
Cyprus and western and southern Anatolia. This alliance
belongs to Juniperetalia haemisphaericae Ri vas-
Martínez &Molina 1999 order distributed
in central and eastern Mediterranean.


Within the Pino-Juniperetea class the pioneer
orophilous vegetation counts, in general, quite a low
number of specialized species. In spite of this, the lack
of character species is compensated by their high biogeographic
significance and allegiance to a precise
ecological context.Pino-Junipereteacommunities ac