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ŠUMARSKI LIST 5-6/2018 str. 39     <-- 39 -->        PDF

individual near Zadar shared its ribotype with Q. crenata from Italy (Latium), while the putative green oak individual from Rijeka exhibited the same ribotype as Q. suber from Croatia (Pula) and Italy (Latium) as well as Q. crenata from Slovenia and Italy (Trentino). Likewise, placement of both Q. × viridis from Zadar and Rijeka as sister taxa to Q. crenata and Q. suber in the MP tree indicates that these taxa (Q. suber, Q. crenata and Q. × viridis) are indeed highly related at the nuclear level. Borzan (2002) already pointed out that “a visit to the Herbarium Patavinum, Padua (Italy), and to the Erbario Centrale Italiano »Filippo Parlatore«, Sezione di Botanica, Museo di Scienze Naturali, Firenze (Italy), revealed that the herbarium samples of the “green oak” from Zadar are very similar to those kept in these Italian botanical institutions under the name Q. crenata and Q. pseudosuber Santi”. Individuals assigned to the complex Q. crenata Q. pseudosuber are in fact traditionally acknowledged as stabilised hybrids between Q. cerris × Q. suber and the latest paper on this topic concluded that Q. crenata Lam. is the correct name for this nothospecies (Cristofolini and Crema 2005; Cristofolini et. al., 2017). This oak taxon (the “false cork oak”) is known to occur in southern France and Italy where the two supposed parental species are found in sympatry; however, scattered Q. crenata individuals can be also found in Northeastern Italy, western Slovenia and Croatia, where Q. suber is currently out of its distribution range (Cristofolini and Crema 2005; Bellarosa et. al., 2005). Thus, the origin of these Q. crenata individuals is still under debate (Cristofolini and Crema, 2005; Conte et. al., 2006; Schirone et. al., 2015).
For example, some authors included Croatia in the natural range of Q. suber (Muller, 1900; see references in Schirone et. al., 2015 and Euro+Med PlantBase) because this oak species has been well documented in older floristic literature from several localities across Croatia. These include Istria and particularly the Šijana forest, near Pula, where the largest and oldest known cork oak stand in Croatia occurs (Trinajstić, 2006 and references therein). In addition, isolated Q. suber trees are known from several localities in